Determined to derail son Preston’s marriage to Irina (guest star Helena Mattsson), Lynette discovers the truth about her future daughter-in-law. In the meantime, Gabrielle offers to help out Bob and Lee in the most generous way, but does not consider the consequences. Bree begins to question whether Sam (guest star Sam Page) has a hidden agenda, and Susan gets creative to help Mike with his finances. Unbeknownst to Angie, Danny meets Patrick Logan (guest star John Barrowman) for the very first time.

Susan and Gabrielle go to extreme and shameless lenghts to top each other’s kids, whereas Bree invites Sam (guest star Sam Page) over for a family dinner. A party is thrown for Mrs. McCluskey after she beats cancer, and Lynette thinks there is more to Preston’s fiancée (guest star Helena Mattsson as Irina) than she’s letting on. Meanwhile, Katherine comes out of the closet, though not willingly.

Gabrielle and Angie head out to New York to track down Ana and Danny, and a chance encounter with supermodels Paulina Porizkova and Heidi Klum, Gaby’s former co-workers, leads the former top model to an unexpected revelation about herself. In the meantime, Mike tries to prove to Susan that he is a man with a capital “M” after he feels emasculated by her, and the Scavo family is surprised to see Preston (former regular castmember Max Carver) back from Europe… accompanied by a hot russian gold digger (guest star Helena Mattson as Irina). Bree discovers a surprising connection to her new employee (guest star Sam Page) and Katherine feels confused regarding her feelings towards Robin (guest star Julie Benz).

Susan causes quite a stir when she invites a former stripper (guest star Julie Benz) into her home, because former exotic dancer Robin will have a profound effect on the ladies of Wisteria Lane: with three teenage boys at home, Lynette will need to keep a watchful eye on them, Bree needs to pick up a few tricks to engage Orson, Katherine needs to find a friendly companion, and Gabrielle needs help with her troublesome niece – and Robin might just provide that help, whether she wants to or not.

Celia comes down with the chicken pox, and having never had it herself, Gabrielle must keep her distance, so she stays with Bob and Lee for a while, and is suddenly reminded of what it feels like to be “single” again. In the meantime, Tom and Lynette forget little Penny’s birthday, which leads the girl to go missing. Bree hires a promising new employee (guest star Sam Page) and Katherine makes a surprising discovery about herself. Meanwhile, Susan cheats on Mrs. McCluskey, whereas Angie grows concerned over Danny’s disappearance.

Gabrielle and Carlos grow concerned with the direction Ana and Danny’s relationship might be taking. In the meantime, Susan befriends a stripper named Robin (guest star Julie Benz) and takes her under her wing, whereas Tom and Lynette come to a disagreement regarding their new therapist (guest star Jane Leeves). Bree watches over Orson as she fears he may commit suicide, and Angie tries to get a neighbor to sort out her garbage, much to Nick’s dismay.

Lynette balks at the idea of seeing a couples’ therapist. In the meantime, Katherine shows no intention to moving back onto the lane, whereas Gabrielle and Susan are determined to find out which of their kids is in the smartest math group at school, whereas Orson makes life harder for Bree and Angie disapproves of Danny’s new girlfriend.

Susan turns up the heat for Mike, whereas Lynette discovers Tom’s true intentions. Gabrielle is forced to examine feelings she’s suppressed regarding her heritage, and Bree learns it’ll be hard to undo the hurt she’s caused Orson. In the meantime, Ana is more determined to land Danny.